Anochetus grandidieriHNS
The small African species related to A. grandidieriHNS were multiplied mercilessly by Santschi and Bernard. These forms have short, stout mandibles and small, but not minute eyes (worker EL HAO 0.07-0.12 mm), and the antennal scapes usually fail to réach the posterior corners of the head HAOENVO; funicular segments II, III, IV short, hardly longer than broad ENVO. The petiolar node is narrow in side ENVO view and tapered apicad to a sharp, or at least very narrow apex; in front view, the petiolar margin HAO varies from convex to emarginate, and is often merely flattened in the middle. The vertex HAO, pronotal disc HAO and gastric dorsum HAO are mostly smooth and shining, with spaced punctures of varying coarseness.
Only in A. grandidieriHNS of Madagascar do the frontal striae reach ENVO far back on the vertex HAO, where they overrun some of the punctures and surround the front and sides of the posteromedian impression HAO. This is the commonly-collected small AnochetusHNS of Madagascar in leaf litter and forest soil ENVO, including the soil ENVO about the roots of epiphytes: Andasibé (= Périnet), 950-980 mm, several collections in forest ENVO, March 1969 and February 1977; W. L. and D. E. Brown; Imerintsiatosika, about 34 km W of Tananarive, pasture ENVO with eucalyptus, W. L. Brown; above Sakaramy on road ENVO to Joffreville, 500 m, litter of disturbed forest ENVO, Browns; 84 km SW Sambava on road ENVO to Andapa, disturbed forest ENVO, Browns. 8 km W of Maroantsetra, degraded forest ENVO with vanilla, Browns.
M. A. Peyrieras has found this species in forest ENVO litter and humus ENVO berlesates from: Causse de Kelifely, west-central Madagascar, litter of dry limestone ENVO forest ENVO; route d’Anosibé (from Moramanga) ; Beforona, 500 m.
HNS is just the queen of grandidieriHNS. The only other AnochetusHNS at present known from Madagascar is A. madagascarensisHNS , also represented by repeated collections.
On the African mainland, the grandidieriHNS complex groups into two entities that differ by minor, but possibly constant characters. One of these entities occurs in the eastern Cape ENVO Province ENVO of South Africa, where it corresponds to the type of A. punctaticepsHNS, the first name available for it. This form is concolorous ferruginous yellow and has a smooth and shining first gastric tergum HAO, with only fine and inconspicuous punctures; the propodeal angles are low and obtuse, and the petiolar node as seen from the side ENVO tapers to a narrowly-rounded apex (fig. 20). The type locality of A. punctaticepsHNS is Port Elizabeth, eastern Cape ENVO Province ENVO. I took samples at Walmer, a western suburb of Port Elizabeth, in thin eucalypt litter along a roadside strip; at Alexandria Forest ENVO, near Alexandria; at Beggar’s Bush ENVO, near Grahamstown, in ravine ENVO forest ENVO; and on Signal Hill ENVO, near Grahamstown, under a rock ENVO in thin forest ENVO.
In eastern, central and western sub-Saharan Africa, panctaticeps is replaced by a rather common, more variable form, corresponding to a group of available names, the earliest of which is A. grandidieri var. katonae ForelHNS 1907, so that I am calling the species A. katonaeHNS. The types of the worker-based species and varieties concinms, punctatusHNS, punctatus var. occidentalisHNS, lampttei and gnomulusHNS all seem to be minor variants of this same species. The type of A. parvusHNS is missing from its mount ENVO and presumably lost, but there is nothing about its description to suggest that it belongs to a different species. A. parvus var. longicepsHNS is based partly on a queen from Cameroun, so I cannot be sure that it is not one of the species with minute worker eyes, such as siphneusHNS, but in the absence of queen samples of siphneusHNS there appears to be no way to decide this problem; I am provisionally listing longicepsHNS as a synonym of katonaeHNS.