Hylodes heyeri Haddad, Pombal & Bastos, 1996

Weber, Luiz Norberto & Caramaschi, Ulisses, 2013, A survey of the internal oral morphology in larvae of the genus Hylodes Fitzinger, 1826 (Amphibia, Anura, Hylodidae), Zootaxa 3635 (5), pp. 557-568: 559

publication ID


publication LSID


persistent identifier


treatment provided by


scientific name

Hylodes heyeri Haddad, Pombal & Bastos, 1996


Hylodes heyeri Haddad, Pombal & Bastos, 1996 

Material: two specimens (stage 28) and one specimen (stage 27).

Ventral aspect ( Fig. 2 View Figure A): Buccal floor triangular, wider than long or equivalent; two pairs of infralabial papillae present, close from one another; the outer pair is longer than wide, has a complex structure with four to five finger-shaped projections of irregular margin that touch each other;, papilla base short, inexistent or less discernible; the inner pair is small, with truncate apex with length slightly lower to the length of the lingual papilla; two filiform lingual papillae present, placed side by side, tapered; lingual papillae closer to the first papilla of the buccal floor arena than to the infralabial papilla; there are about 20 papillae on each side, enclosing the buccal floor arena and extending from the region near the tongue rudiments to near the ventral velum; papillae directed to the arena center, finger-shaped and curved; the bigger papilla of the floor arena is bifurcate, located in the middle portion of the arena; some arena papillae show irregular margin; some small and conical papillae located at the base of arena; about three to four prepocket papillae, small to medium-sized, similar to those from the buccal floor arena; large number of pustules present on the arena surface, more concentrated in its posterior portion; ventral velum with accentuated median notch, undulated margin and about six finger-shaped projections, developed, above the glottis.

Dorsal aspect ( Fig. 2 View Figure B): Buccal roof triangular, somewhat shaped as the floor; prenarial arena with low crest and shaped as a reversed V; reniform choanae, separated and oriented at an angle of 35 ° in relation to the transverse plane; distance between choanae about one-third less than to the width of the median-ridge; pre-choanal papillae not found; postnarial arena with five to seven papillae, arranged on each side in a separate row, papillae with varied sizes and finger-shaped; largest papillae postero-located, finger-shaped, with sharp apex, irregular antero-margin and directed to the center; cluster of papillae in postnarial arena forming an inverted V; one to three small papillae on the same transverse plane, located in front of the median ridge; complex lateral-ridge papillae, hand-shaped with five to six ramifications of anterior margin finely serrate and directed to the center; the median ridge half circleshaped with irregular margin; buccal roof arena bordered by 11–22 papillae on each side, finger-shaped, tapered, and with anterior margin aliasing, directed up and to the arena center; absence of bifurcate papilla in the arena; arena papillae forming an approximate U design; pustules are present in large number and evenly distributed; distinct row with about eight lateral papillae on the roof arena; distinct glandular area; dorsal velum medially interrupted, having about six to eight uniform projections of medium size.